diff --git a/app/models/balance/base_calculator.rb b/app/models/balance/base_calculator.rb
index 0e4b5d24a8..319362e0d2 100644
--- a/app/models/balance/base_calculator.rb
+++ b/app/models/balance/base_calculator.rb
@@ -66,6 +66,11 @@ def non_cash_adjustments_for_date(start_non_cash, end_non_cash, non_cash_flows)
       end_non_cash - start_non_cash - non_cash_flows
     end
 
+    # Keeps asset/liability flow sign conventions centralized for persisted balances.
+    def flows_factor
+      account.classification == "asset" ? 1 : -1
+    end
+
     # If holdings value goes from $100 -> $200 (change_holdings_value is $100)
     # And non-cash flows (i.e. "buys") for day are +$50 (net_buy_sell_value is $50)
     # That means value increased by $100, where $50 of that is due to the change in holdings value, and $50 is due to the buy/sell
@@ -159,7 +164,7 @@ def build_balance(date:, **args)
         cash_adjustments: args[:cash_adjustments] || 0,
         non_cash_adjustments: args[:non_cash_adjustments] || 0,
         net_market_flows: args[:net_market_flows] || 0,
-        flows_factor: account.classification == "asset" ? 1 : -1
+        flows_factor: flows_factor
       )
     end
 end
diff --git a/app/models/balance/forward_calculator.rb b/app/models/balance/forward_calculator.rb
index 29163b7c6e..521473693c 100644
--- a/app/models/balance/forward_calculator.rb
+++ b/app/models/balance/forward_calculator.rb
@@ -156,8 +156,4 @@ def derive_end_cash_balance(start_cash_balance:, date:)
     def derive_end_non_cash_balance(start_non_cash_balance:, date:)
       derive_non_cash_balance(start_non_cash_balance, date, direction: :forward)
     end
-
-    def flows_factor
-      account.asset? ? 1 : -1
-    end
 end
diff --git a/app/models/balance/reverse_calculator.rb b/app/models/balance/reverse_calculator.rb
index d073beda74..64e9993757 100644
--- a/app/models/balance/reverse_calculator.rb
+++ b/app/models/balance/reverse_calculator.rb
@@ -18,6 +18,8 @@ def calculate
       account.current_anchor_date.downto(calculation_start_date).map do |date|
         flows = flows_for_date(date)
         valuation = sync_cache.get_valuation(date)
+        cash_adjustments = 0
+        non_cash_adjustments = 0
 
         if use_opening_anchor_for_date?(date)
           end_cash_balance = derive_cash_balance_on_date_from_total(
@@ -50,6 +52,20 @@ def calculate
           market_value_change = market_value_change_on_date(date, flows)
         end
 
+        if use_opening_boundary_adjustment_for_date?(date)
+          boundary_adjustment = opening_boundary_adjustment(
+            end_cash_balance: end_cash_balance,
+            end_non_cash_balance: end_non_cash_balance,
+            flows: flows,
+            market_value_change: market_value_change
+          )
+
+          start_cash_balance = boundary_adjustment[:start_cash_balance]
+          start_non_cash_balance = boundary_adjustment[:start_non_cash_balance]
+          cash_adjustments = boundary_adjustment[:cash_adjustments]
+          non_cash_adjustments = boundary_adjustment[:non_cash_adjustments]
+        end
+
         output_balance = build_balance(
           date: date,
           balance: end_cash_balance + end_non_cash_balance,
@@ -60,6 +76,8 @@ def calculate
           cash_outflows: flows[:cash_outflows],
           non_cash_inflows: flows[:non_cash_inflows],
           non_cash_outflows: flows[:non_cash_outflows],
+          cash_adjustments: cash_adjustments,
+          non_cash_adjustments: non_cash_adjustments,
           net_market_flows: market_value_change
         )
 
@@ -100,4 +118,51 @@ def derive_start_non_cash_balance(end_non_cash_balance:, date:)
     def use_opening_anchor_for_date?(date)
       account.has_opening_anchor? && date == account.opening_anchor_date
     end
+
+    # Applies the one-day bridge from the opening anchor to the first derived day.
+    def use_opening_boundary_adjustment_for_date?(date)
+      account.has_opening_anchor? && date == account.opening_anchor_date.next_day
+    end
+
+    # Builds explicit adjustments that make the opening-boundary row auditably reconcile.
+    def opening_boundary_adjustment(end_cash_balance:, end_non_cash_balance:, flows:, market_value_change:)
+      opening_cash_balance, opening_non_cash_balance = opening_balance_components
+
+      {
+        start_cash_balance: opening_cash_balance,
+        start_non_cash_balance: opening_non_cash_balance,
+        cash_adjustments: cash_adjustments_for_date(opening_cash_balance, end_cash_balance, cash_flows_total(flows)),
+        non_cash_adjustments: opening_boundary_non_cash_adjustments(
+          opening_non_cash_balance: opening_non_cash_balance,
+          end_non_cash_balance: end_non_cash_balance,
+          flows: flows,
+          market_value_change: market_value_change
+        )
+      }
+    end
+
+    # Splits the opening anchor total into the calculator's persisted components.
+    def opening_balance_components
+      opening_cash_balance = derive_cash_balance_on_date_from_total(
+        total_balance: account.opening_anchor_balance,
+        date: account.opening_anchor_date
+      )
+
+      [ opening_cash_balance, account.opening_anchor_balance - opening_cash_balance ]
+    end
+
+    # Converts same-day cash flow columns into their signed balance impact.
+    def cash_flows_total(flows)
+      (flows[:cash_inflows] - flows[:cash_outflows]) * flows_factor
+    end
+
+    # Converts same-day non-cash flow columns into their signed balance impact.
+    def non_cash_flows_total(flows)
+      (flows[:non_cash_inflows] - flows[:non_cash_outflows]) * flows_factor
+    end
+
+    # Keeps boundary non-cash math market-value-aware for investment accounts.
+    def opening_boundary_non_cash_adjustments(opening_non_cash_balance:, end_non_cash_balance:, flows:, market_value_change:)
+      end_non_cash_balance - opening_non_cash_balance - non_cash_flows_total(flows) - market_value_change
+    end
 end
diff --git a/test/models/balance/materializer_test.rb b/test/models/balance/materializer_test.rb
index 6ee91d99db..741260601d 100644
--- a/test/models/balance/materializer_test.rb
+++ b/test/models/balance/materializer_test.rb
@@ -268,6 +268,59 @@ class Balance::MaterializerTest < ActiveSupport::TestCase
     assert_balance_fields_persisted(expected_balances)
   end
 
+  test "reverse materialization persists opening boundary adjustment" do
+    account = families(:empty).accounts.create!(
+      name: "Linked Depository",
+      balance: 1000,
+      cash_balance: 1000,
+      currency: "USD",
+      accountable: Depository.new
+    )
+    opening_date = Date.new(2024, 1, 1)
+    boundary_date = opening_date + 1.day
+    transaction_date = opening_date + 2.days
+    current_anchor_date = opening_date + 3.days
+
+    account.entries.create!(
+      name: "Current Balance",
+      date: current_anchor_date,
+      amount: 1000,
+      currency: "USD",
+      entryable: Valuation.new(kind: "current_anchor")
+    )
+    account.entries.create!(
+      name: "Transaction",
+      date: transaction_date,
+      amount: 200,
+      currency: "USD",
+      entryable: Transaction.new
+    )
+    account.entries.create!(
+      name: "Opening Balance",
+      date: opening_date,
+      amount: 1000,
+      currency: "USD",
+      entryable: Valuation.new(kind: "opening_anchor")
+    )
+
+    Holding::Materializer.any_instance.expects(:materialize_holdings).returns([]).once
+
+    Balance::Materializer.new(account, strategy: :reverse).materialize_balances
+
+    opening_balance = account.balances.find_by!(date: opening_date)
+    boundary_balance = account.balances.find_by!(date: boundary_date)
+    transaction_balance = account.balances.find_by!(date: transaction_date)
+
+    assert_equal 1000, opening_balance.end_balance
+    assert_equal 1000, boundary_balance.start_balance
+    assert_equal 1200, boundary_balance.end_balance
+    assert_equal 200, boundary_balance.cash_adjustments
+    assert_equal 0, boundary_balance.cash_inflows
+    assert_equal 0, boundary_balance.cash_outflows
+    assert_equal 1200, transaction_balance.start_balance
+    assert_equal 1000, transaction_balance.end_balance
+  end
+
   private
 
     def assert_balance_fields_persisted(expected_balances)
diff --git a/test/models/balance/reverse_calculator_test.rb b/test/models/balance/reverse_calculator_test.rb
index 070e1f0ff0..f40a134b1a 100644
--- a/test/models/balance/reverse_calculator_test.rb
+++ b/test/models/balance/reverse_calculator_test.rb
@@ -28,6 +28,102 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
     )
   end
 
+  test "opening boundary difference is surfaced as an adjustment instead of a silent jump" do
+    opening_date = Date.new(2024, 1, 1)
+    day_after_opening = opening_date + 1.day
+    transaction_date = opening_date + 2.days
+    current_anchor_date = transaction_date + 1.day
+
+    account = create_account_with_ledger(
+      account: { type: Depository, balance: 1000, cash_balance: 1000, currency: "USD" },
+      entries: [
+        { type: "current_anchor", date: current_anchor_date, balance: 1000 },
+        { type: "transaction", date: transaction_date, amount: 200 },
+        { type: "opening_anchor", date: opening_date, balance: 1000 }
+      ]
+    )
+
+    calculated = Balance::ReverseCalculator.new(account).calculate
+
+    assert_calculated_ledger_balances(
+      calculated_data: calculated,
+      expected_data: [
+        {
+          date: opening_date,
+          legacy_balances: { balance: 1000, cash_balance: 1000 },
+          balances: { start: 1000, start_cash: 1000, start_non_cash: 0, end_cash: 1000, end_non_cash: 0, end: 1000 },
+          flows: 0,
+          adjustments: 0
+        },
+        {
+          date: day_after_opening,
+          legacy_balances: { balance: 1200, cash_balance: 1200 },
+          balances: { start: 1000, start_cash: 1000, start_non_cash: 0, end_cash: 1200, end_non_cash: 0, end: 1200 },
+          flows: 0,
+          adjustments: { cash_adjustments: 200, non_cash_adjustments: 0 }
+        },
+        {
+          date: transaction_date,
+          legacy_balances: { balance: 1000, cash_balance: 1000 },
+          balances: { start: 1200, start_cash: 1200, start_non_cash: 0, end_cash: 1000, end_non_cash: 0, end: 1000 },
+          flows: { cash_inflows: 0, cash_outflows: 200 },
+          adjustments: 0
+        },
+        {
+          date: current_anchor_date,
+          legacy_balances: { balance: 1000, cash_balance: 1000 },
+          balances: { start: 1000, start_cash: 1000, start_non_cash: 0, end_cash: 1000, end_non_cash: 0, end: 1000 },
+          flows: 0,
+          adjustments: 0
+        }
+      ]
+    )
+  end
+
+  test "opening boundary adjustment uses liability flow direction" do
+    opening_date = Date.new(2024, 1, 1)
+    boundary_date = opening_date + 1.day
+    current_anchor_date = boundary_date + 1.day
+
+    account = create_account_with_ledger(
+      account: { type: CreditCard, balance: 500, cash_balance: 500, currency: "USD" },
+      entries: [
+        { type: "current_anchor", date: current_anchor_date, balance: 500 },
+        { type: "transaction", date: boundary_date, amount: 100 },
+        { type: "opening_anchor", date: opening_date, balance: 1000 }
+      ]
+    )
+
+    calculated = Balance::ReverseCalculator.new(account).calculate
+
+    assert_calculated_ledger_balances(
+      calculated_data: calculated,
+      expected_data: [
+        {
+          date: opening_date,
+          legacy_balances: { balance: 1000, cash_balance: 1000 },
+          balances: { start: 1000, start_cash: 1000, start_non_cash: 0, end_cash: 1000, end_non_cash: 0, end: 1000 },
+          flows: 0,
+          adjustments: 0
+        },
+        {
+          date: boundary_date,
+          legacy_balances: { balance: 500, cash_balance: 500 },
+          balances: { start: 1000, start_cash: 1000, start_non_cash: 0, end_cash: 500, end_non_cash: 0, end: 500 },
+          flows: { cash_inflows: 0, cash_outflows: 100 },
+          adjustments: { cash_adjustments: -600, non_cash_adjustments: 0 }
+        },
+        {
+          date: current_anchor_date,
+          legacy_balances: { balance: 500, cash_balance: 500 },
+          balances: { start: 500, start_cash: 500, start_non_cash: 0, end_cash: 500, end_non_cash: 0, end: 500 },
+          flows: 0,
+          adjustments: 0
+        }
+      ]
+    )
+  end
+
   # Reconciliation valuations act as waypoints during reverse syncs. This ensures that
   # historical balances accurately reflect the API-reported values, even if the transaction
   # history is incomplete or missing.
@@ -43,6 +139,8 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
 
     calculated = Balance::ReverseCalculator.new(account).calculate
 
+    # The day after the opening anchor now carries an explicit adjustment so
+    # the gap to the first reconciliation waypoint is auditable.
     assert_calculated_ledger_balances(
       calculated_data: calculated,
       expected_data: [
@@ -70,10 +168,10 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
         {
           date: 3.days.ago,
           legacy_balances: { balance: 17000, cash_balance: 17000 },
-          balances: { start: 17000, start_cash: 17000, start_non_cash: 0, end_cash: 17000, end_non_cash: 0, end: 17000 },
+          balances: { start: 15000, start_cash: 15000, start_non_cash: 0, end_cash: 17000, end_non_cash: 0, end: 17000 },
           flows: 0,
-          adjustments: { cash_adjustments: 0, non_cash_adjustments: 0 }
-        }, # Derived from Reconciliation waypoint
+          adjustments: { cash_adjustments: 2000, non_cash_adjustments: 0 }
+        }, # Opening boundary adjustment explains the gap from opening anchor to waypoint
         {
           date: 4.days.ago,
           legacy_balances: { balance: 15000, cash_balance: 15000 },
@@ -154,10 +252,10 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
         {
           date: 6.days.ago,
           legacy_balances: { balance: 22500, cash_balance: 22500 },
-          balances: { start: 22200, start_cash: 22200, start_non_cash: 0, end_cash: 22500, end_non_cash: 0, end: 22500 },
+          balances: { start: 18000, start_cash: 18000, start_non_cash: 0, end_cash: 22500, end_non_cash: 0, end: 22500 },
           flows: { cash_inflows: 300, cash_outflows: 0 },
-          adjustments: { cash_adjustments: 0, non_cash_adjustments: 0 }
-        }, # Income derived further back, right before opening_anchor
+          adjustments: { cash_adjustments: 4200, non_cash_adjustments: 0 }
+        }, # Opening boundary adjustment explains the gap from opening anchor to derived balance
         {
           date: 7.days.ago,
           legacy_balances: { balance: 18000, cash_balance: 18000 },
@@ -216,9 +314,9 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
         {
           date: 3.days.ago,
           legacy_balances: { balance: 18000, cash_balance: 18000 },
-          balances: { start: 18000, start_cash: 18000, start_non_cash: 0, end_cash: 18000, end_non_cash: 0, end: 18000 },
+          balances: { start: 15000, start_cash: 15000, start_non_cash: 0, end_cash: 18000, end_non_cash: 0, end: 18000 },
           flows: 0,
-          adjustments: { cash_adjustments: 0, non_cash_adjustments: 0 }
+          adjustments: { cash_adjustments: 3000, non_cash_adjustments: 0 }
         },
         {
           date: 4.days.ago,
@@ -314,10 +412,10 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
         {
           date: Date.current,
           legacy_balances: { balance: 20000, cash_balance: 10000 },
-          balances: { start: 20000, start_cash: 10000, start_non_cash: 10000, end_cash: 10000, end_non_cash: 10000, end: 20000 },
+          balances: { start: 15000, start_cash: 5000, start_non_cash: 10000, end_cash: 10000, end_non_cash: 10000, end: 20000 },
           flows: { market_flows: 0 },
-          adjustments: 0
-        }, # Since $10,000 of holdings, cash has to be $10,000 to reach $20,000 total value
+          adjustments: { cash_adjustments: 5000, non_cash_adjustments: 0 }
+        }, # Opening boundary adjustment explains the gap to the current provider anchor
         {
           date: 1.day.ago,
           legacy_balances: { balance: 15000, cash_balance: 5000 },
@@ -672,9 +770,9 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
         {
           date: 1.day.ago,
           legacy_balances: { balance: 20000, cash_balance: 19000 },
-          balances: { start: 20000, start_cash: 19000, start_non_cash: 1000, end_cash: 19000, end_non_cash: 1000, end: 20000 },
+          balances: { start: 15000, start_cash: 14000, start_non_cash: 1000, end_cash: 19000, end_non_cash: 1000, end: 20000 },
           flows: { market_flows: 0 },
-          adjustments: 0
+          adjustments: { cash_adjustments: 5000, non_cash_adjustments: 0 }
         },
         {
           date: 2.days.ago,
@@ -719,10 +817,10 @@ class Balance::ReverseCalculatorTest < ActiveSupport::TestCase
         {
           date: 1.day.ago,
           legacy_balances: { balance: 20000, cash_balance: 20000 },
-          balances: { start: 20000, start_cash: 20000, start_non_cash: 0, end_cash: 20000, end_non_cash: 0, end: 20000 },
+          balances: { start: 15000, start_cash: 15000, start_non_cash: 0, end_cash: 20000, end_non_cash: 0, end: 20000 },
           flows: 0,
-          adjustments: 0
-        }, # Gap above the anchor carries down with no flows
+          adjustments: { cash_adjustments: 5000, non_cash_adjustments: 0 }
+        }, # Opening boundary adjustment explains the gap above the anchor
         {
           date: 2.days.ago,
           legacy_balances: { balance: 15000, cash_balance: 15000 },